13 | | where ''P'',,''i'',, is the total production rate of (''i''), ~~Y''''i'''' is the total fishery catch rate of (''i''), M2''i'' is the total predation rate for group (''i''), ''B'',,i,, the biomass of the group, E''i'' the net migration rate (emigration – immigration), ''BA'',,''i'',, is the biomass accumulation rate for (''i''), while ''M0'',,''i'',,'' = P'',,''i'',, · (1-EE''i''~~) is the 'other mortality' rate for (''i''). |

| 13 | where ''P'',,''i'',, is the total production rate of (''i''), ''Y,,i,,'' is the total fishery catch rate of (''i''), ''M2,,i,,'' is the total predation rate for group (''i''), ''B,,i,,'' the biomass of the group, E,,i,, the net migration rate (emigration – immigration), ''BA'',,''i'',, is the biomass accumulation rate for (''i''), while ''M0,,i'',,'' = ''P,,i'',, . (1-EE,,i,,) is the 'other mortality' rate for (''i''). |